I think straight-pull spokes don't have a problem with negative loads the same way j-bend spokes do.
For j-bend spokes, any push-force incurred by rim flex would force a non-interlaced spoke into substantial buckling and twisting. It is substantial because if the spoke started out perfectly straight, there is initially no direction it could move to take up any approach of the nipple towards the hub, the spoke first needs to deform into a new buckle that can then start to take up the slack. Juhist's answer discusses this in some more mathematical detail.
It is this motion that causes the nipples to loosen. Interlacing "solves" the problem by forcing every spoke into a little pre-buckle, which gives them a well-defined direction into which to deform when slack needs to be taken up. It's a big ugly hack, but it mostly avoids twisting that would unscrew the nipple.
Straigh-pull spokes solve the problem in a much better way: they allow the spokes to simply move inwards at the hub when necessary (in a way the j-bend prevents, counter-productively). Movement at the hub is harmless as there's no screwed thread at that end.
As a secondary advantage, the perfectly straight spokes make the wheel stiffer by way of being more rigid in the tensile direction, which prevents the wheel from flexing as much in the first place.